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Tom Kaye, Ph.D.


Tom Kaye

Tom Kaye is Executive Director of the Institute for Applied Ecology. He graduated with a B.S. from The Evergreen State College (1984) and received a master's degree (1989) and PhD (2001) from Oregon State University. After working for Olympic National Park (1984-1987), he joined the Oregon Department of Agriculture's Plant Conservation Biology Program where he conducted research and contributed to policy on management of threatened and endangered plant species. He has served on the IUCN Species Survival Commission, Re-introduction Specialist Group and is a member of the Native Plant Society of Oregon.

Tom specializes in endangered species propagation and restoration, the population dynamics of rare plants, population viability analysis, and monitoring. In addition, his interests include plant-pollinator interactions and plant systematics. He has a long-standing interest in coastal vegetation, including general patterns of plant distributions on the Pacific Coast and restoration of endangered species of beaches and dunes.

As Assistant Professor (courtesy) in the Department of Botany and Plant Pathology at Oregon State University, Tom supervises graduate students, serves on graduate student committees, present guest lectures, and leads discussion groups in a variety of ecological topics.

Email Tom

(541) 753-3099 ext. 111

Selected publications (to request reprints, please email Tom):

Richardson, P.J., A.S. MacDougall, A.G. Stanley, T.N. Kaye, and P.W. Dunwiddie.  In press.  Inversion of dominance-diversity relationships along a latitudinal stress gradient.  Ecology.

Ellis, M.M., J.L. Williams, P. Lesica, T.J. Bell, P. Bierzychudek, M. Bowles, E. Crone, D.F. Doak, J. Ehrlen, A. Ellis-Adam, K. McEachern, R. Ganesan, P. Latham, S. Luijten, T.N. Kaye, T.M. Knight, E.S. Menges, W.F. Morris, H. den Nijs, G. Oostermeijer, P.F. Quintana-Ascencio, J.S. Shelley, A. Stanley, A. Thorpe, T. Ticktin, T. Valverde, and C. Weekley. In press.  Matrix population models from 20 studies of perennial plant populations.  Ecology.

Seabloom, E.W., C.D. Benfield, E.T. Borer, A.G. Stanley, T.N. Kaye, and P.W. Dunwiddie.  2011.  Provenance, lifespan, and phylogeny do not affect grass species’ responses to nitrogen and phosphorus.  Ecological Applications 21:2129-2142.

Thorpe, A.S. and T.N. Kaye.  2011.  Conservation and reintroduction of the endangered Willamette daisy.  Native Plants Journal 12:289-298.

Stanley, A.G., P.W. Dunwiddie, and T.N. Kaye.  2011.  Restoring invaded Pacific Northwest prairies: management recommendations from a region-wide experiment. Northwest Science 85:233-246. 

Dennehy, C., H.E. Anderson, D.R. Clements, E.R. Alverson, T.N. Kaye, and R. Gilbert.  2011.  Management strategies for invasive plants in Pacific Northwest prairies and oak woodlands.  Northwest Science 85:329-351.

Kaye, T.N.  2011.  Toward successful reintroductions: The combined importance of species traits, site quality, and restoration technique. In, J.W. Willoughby, B.K. Orr, K.A. Schierenbeck, and N.J. Jensen [eds.], Proceedings of the CNPS Conservation Conference: Strategies and Solutions, 17-19 Jan 2009, California Native Plant Society, Sacramento, CA. pp. 99-106.

Kaye, T.N., R. Schwindt, and C. Menke.  2011.  Do private land owners support species conservation?  Results of a local survey (Oregon).  Ecological Restoration 29:10-12.

Stanley, AG., Thomas N. Kaye, and Peter Dunwiddie.  2011.  Multiple treatment combinations and seed addition increase abundance and diversity of native plants in Pacific Northwest prairies.  Ecological Restoration 29:35-44.

Crone, E., E. Menges, M. Ellis, T. Bell, P. Bierzychudek, J. Ehrlen, T.N. Kaye, T. Knight, P. Lesica, W. Morris, G. Oostermeijer, P. Quintana-Ascencio, A. Stanley, T. Valverde, T. Ticktin, and J. Williams. 2011. How do plant ecologists use matrix population models?  Ecology Letters 14:1-8.  doi: 10.1111/j.1461-0248.2010.01540.x

Miller, S.A., A. Bartow, M. Gisler, K. Ward, A.S. Young, and T.N. Kaye. (corresponding author)  2011.  Can an ecoregion serve as a seed transfer zone? Evidence from a common garden study with five native species.  Restoration Ecology 19:268-276.  doi: 10.1111/j.1526-100X.2010.00702.x

Wilson, B.L., V. Hipkins, and T.N. Kaye.  2010.  One taxon or two: Are Frasera umpquaensis and F. fastigiata (Gentianaceae) distinct species?  Madrono 57:106-119.

Lawrence, B.A. and T.N. Kaye.  2009.  Reintroduction of Castilleja levisecta: Effects of ecological similarity, source population genetics, and habitat quality.  Restoration Ecology.  doi: 10.1111/j.1526-100X.2009.00549.x

Kaye, T.N.  2008.  Vital steps toward success of endangered plant reintroductions.  Native Plants Journal 9:313-322.

Stanley, A.S., T.N. Kaye, and P.W. Dunwiddie.  2008.  Regional strategies for restoring invaded prairies: observations from a multisite collaborative research project.  Native Plants Journal 9: 247-254.

Lawrence, B.A. and T.N. Kaye.  2008.  Direct and indirect effects of host plants: Implications for reintroduction of an endangered hemiparasitic plant (Castilleja levisecta).  Madroño 55:151-158.

Kaye, T.N., M. Blakeley-Smith, and W. Thies. 2008.  Long term effects of post-harvest stump removal and N-fertilization on understory vegetation in western USA forests.  Forest Ecology and Management 256:732-740.

Meinke, R.J. and T.N. Kaye.  2007.  Kalmiopsis fragrans (Ericaceae), a new distylous species from the southern Cascade Mountains of Oregon.  Journal of the Botanical Research Institute of Texas (formerly SIDA).  Volume 1, No.1.

Guerrant, E.O. and T.N. Kaye.  2007.  Reintroduction of rare and endangered plants: Common factors, questions and approaches.  Australian Journal of Botany 55:362-270.

Kaye, T.N. and M. Blakeley-Smith.  2006.  False-brome (Brachypodium sylvaticum).  In, Invasive Species in the Pacific Northwest. P.D. Boersma, S.E. Reichard, and A.N. van Buren, editors.  University of Washington Press, Seattle. Pages 80-81.

Lawrence, B.A. and T.N. Kaye.  2006. Habitat variation throughout the historic range of golden paintbrush, a Pacific Northwest prairie endemic: Implications for reintroduction.  Northwest Science 80:140-152.

Lawrence, B.A. and T.N. Kaye.  2005.  Growing Castilleja for restoration and the garden.  Rock Garden Quarterly 63:128-134.

Wilson, B.L., V.D. Hipkins, E. Rey-Vizgirdas, and T.N. Kaye.  2005.  Variation in Lewisia kelloggii (Portulacaceae) with description of a new species endemic to Idaho.  Western North American Naturalist 65:345-358.

Kaye, T.N.  2004.  Reintroducing the endangered pink sand-verbena to Pacific Coast beaches: Direct seeding and out-planting.  In, Brooks, M.B., S.K. Carothers, and T. LaBlanca, eds.  The Ecology and Management of Rare Plants of Northwestern California: Proceedings from a 2002 Symposium of the North Coast Chapter of the California Native Plant Society.  California Native Plant Society, Sacramento, California.

Wilson, M.V, T. Erhart, P.C. Hammond, T.N. Kaye, K. Kuykendall, A. Liston, A.F. Robinson, Jr., C.B. Schultz, and P.M. Severns.  2003.  Biology of Kincaid’s lupine (Lupinus sulphureus ssp. kincaidii [Smith] Phillips), a threatened species of western Oregon native prairies.  Natural Areas Journal 23:72-83.

Kaye, T.N. and D.A. Pyke.  2003.  The effect of stochastic technique on estimates of population viability from transition matrix models.  Ecology 1464-1476.

Kaye, T.N. and J. Cramer.  2003.  Direct seeding or transplanting: The cost of restoring populations of Kincaid's lupine (Oregon).  Ecological Restoration 21:224-225.

McGlaughlin, M., K. Karoly, and T.N. Kaye.  2002.  Genetic variation and its relationship to population size in reintroduced populations of the pink sand verbena, Abronia umbellata subsp. breviflora.  Conservation Genetics 3:411-420.

Kaye, T.N., K. Pendergrass, K. Findley and J.B. Kauffman.  2001.  The effect of fire on the population viability of an endangered prairie plant.  Ecological Applications 11:1366-1380.

Caswell, H. and T.N. Kaye.  2001.  Stochastic demography and conservation of an endangered perennial plant (Lomatium bradshawii) in a dynamic fire regime.  Advances in Ecological Research 32:1-51.

Kaye, T.N. and K. Kuykendall.  2001.  Effects of scarification and cold stratification on germination of Lupinus sulphureus ssp. kincaidii.  Seed Science and Technology 29:663-668.

Kaye, T.N.  2001.  Common ground and controversy in native plant restoration: the SOMS debate, source distance, plant selections, and a restoration-oriented definition of native.  In, Native Plant Propagation and Restoration Strategies.  R. Rose and D. Haase, editors.  Nursery Technology Cooperative and Western Forestry and Conservation Association, Corvallis, Oregon.

            Reprinted in: Grasslands 17:4-9, 2007, and Fourth Corner Nurseries catalog, 1-15, 2007.

Kaye, T.N. and K. Kuykendall.  2001.  Germination and propagation techniques for restoring rare Pacific Northwest prairie plants.  In, Reichard, S.H., P.W. Dunwiddie, J.G. Gamon, A.R. Kruckeberg, D.L. Salstrom, editors.  Conservation of Washington’s Native Plants and Ecosystems.  Washington Native Plant Society, Seattle, Washington.

Kaye, T.N.  1999.  Propagation of endangered species: Variable germination of pink sandverbena from Pacific Coast beaches.  Combined Proceedings of the International Plant Propagators Society 49:617-621.

Kaye, T.N., M. Kirkland, and N. Testa.  1999.  Growing endangered plants to save them: germinating, propagating and restoring pink sandverbena.  In, Native plants: propagating and planting.  R. Rose and D. Haase, editors.  Forest Research Lab, Oregon State University.  Pp. 100-107.

Kaye, T.N. and M. Kirkland.  1999.  Effect of timber harvest on Cimicifuga elata, a rare plant of western forests.  Northwest Science 73:159-167.

Kaye, T.N.  1999.  Obligate insect pollination of a rare plant: Lupinus sulphureus ssp. kincaidii.  Northwest Science 73:50-52.

Kaye, T.N.  1999.  From flowering to dispersal: reproductive ecology of an endemic plant, Astragalus australis var. olympicus (Fabaceae).  American Journal of Botany 86:1248-1256.

Pendergrass, K.L., P.M. Miller, J.B. Kauffman, and T.N. Kaye.  1999.  The role of prescribed burning in maintenance of an endangered plant species, Lomatium bradshawii.  Ecological Applications 9:1420-1429.

Kaye, T.N., K. Amsberry, S.D. Gisler, and R.J. Meinke.  1998.  Back from the brink: the challenges of endangered plant re-introduction.  Hortus West 9:4-9, 90-91.

Kaye, T.N., A. Liston, R.M. Love, D. Luoma, R.J. Meinke, and M.V. Wilson [editors].  1997.  Conservation and management of native plants and fungi.  Native Plant Society of Oregon, Corvallis, Oregon.

Kaye, T.N., R.J. Meinke, J. Kagan, S. Vrilakas, K.L. Chambers, P.F. Zika, and J.K. Nelson.  1997.  Patterns of rarity in the Oregon flora: implications for conservation and management.  In, Kaye, T.N., A. Liston, R.M. Love, D. Luoma, R.J. Meinke, and M.V. Wilson [editors].   Conservation and management of native plants and fungi.  Native Plant Society of Oregon, Corvallis, Oregon.

Kaye, T.N.  1997.  Seed dormancy in high elevation plants: implications for ecology and restoration.  In, Kaye, T.N., A. Liston, R.M. Love, D. Luoma, R.J. Meinke, and M.V. Wilson [editors].   Conservation and management of native plants and fungi.  Native Plant Society of Oregon, Corvallis, Oregon.

Greenlee, J. and T.N. Kaye.  1997.  Stochastic matrix projection: a comparison of the effect of element and matrix selection methods on quasi-extinction risk for the rare plant Haplopappus radiatus.  In, Kaye, T.N., A. Liston, R.M. Love, D. Luoma, R.J. Meinke, and M.V. Wilson [editors].   Conservation and management of native plants and fungi.  Native Plant Society of Oregon, Corvallis, Oregon.

Buckingham, Nelsa, E.G. Schreiner, T.N. Kaye, and J.E. Burger, and E.L. Tisch.  1995.  Flora of the Olympic Peninsula.  Northwest Interpretive Association and the Washington Native Plant Society.  Seattle, Washington.  199 pp.

Schreiner, E.G., M.B. Gracz, T.N. Kaye, M.B. Woodward, and N.M. Buckingham.  1994.  Chapter 12, Rare plants. In, D.B. Houston, E.G. Schreiner, and B.B. Moorhead, editors.  Mountain goats in Olympic National Park: biology and management of an introduced species.  Scientific Monograph NPS/NROLYM/NRSM-94/25, United States Department of the Interior, National Park Service.  Pp. 173-185.

Kaye, T.N.  1994.  Cimicifuga elata (tall bugbane) conservation and management.  Kalmiopsis 4:28-29.

Kaye, T.N.  1992.  Bradshaw's desert parsley:  population monitoring and pollination biology.  Kalmiopsis 2:1-4.

Meinke, R.J. and T.N. Kaye.  1992.  Taxonomic assessment of Astragalus tegetarioides (Fabaceae) and a new related species from northern California.  Madroño 39:193-204.

Kaye, T.N. and E. McEvoy.  1992.  Native Plant Society of Oregon policy statement on reintroduction of extirpated or rare species.  Bulletin of the Native Plant Society of Oregon 25:5-6.

Kaye, T.N.  1991.  Inventory of the vascular flora of Dungeness Spit, Clallam County, Washington.  Douglasia Occasional Papers 4:2-25.

Kaye, T.N.  1991.  Chromosome numbers for Eriogonum crosbyae, E. prociduum, Ivesia baileyi var. beneolens, and I. rhypara var. shellyi.  IOPB Newsletter 17, Chromosome Data section.

Kaye, T.N.  1990.  Newberry's gentian and the effect of biased collections on taxonomy.  Bulletin of the Native Plant Society of Oregon 24:131.

Kaye, T.N.  1990.  Chromosome numbers for Crepis nana ssp. ramosa, Senecio flettii (Asteraceae), Campanula piperi (Campanulaceae), Astragalus australis var. olympicus (Fabaceae), Petrophytum hendersonii (Rosaceae), and Pedicularis bracteosa var. atrosanguinea (Scrophulariaceae).  IOPB Newsletter 15, Chromosome Data section.

Kaye, T.N.  1989.  Rare plant field report: Senecio ertterae in Leslie Gulch.  Bulletin of the Native Plant Society of Oregon 22:122-124.

Kaye, T.N.  1988.  Endemism and rarity in plants.  Bulletin of the Native Plant Society of Oregon 22:23-24.


Papers and books published on the internet (non-refereed)

Kaye, T.N., M. Gisler, and R. Fiegener (editors).  2003.  Proceedings of a Conference on Native Plant Restoration and Management on Public Lands in the Pacific Northwest: Rare Plants, Invasive Species and Ecosystem Management. Institute for Applied Ecology, Corvallis, Oregon (on-line abstracts).        


Kaye, T.N.  2001.  Brachypodium sylvaticum (Poaceae) in the Pacific Northwest.  Botanical Electronic News No. 277.  Available on the web at:

Kaye, T.N.  2000.  Notes on the geographic distribution and taxonomy of Lupinus sulphureus ssp. kincaidii (Fabaceae): Watch for it in your neighborhood.  Botanical Electronic News No. 243.  Available on the web at:

Kaye, T.N.  1995.  Cimicifuga elata - some comments on its ecology & distribution.  Botanical Electronic News No. 121.  Available on the web at:


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